1.2 Treatment of Taxa in the Most Recent Revision of Actinidia ........16
1.2.1 Three Varieties of Actinidia arguta (Siebold & Zuccarini) Planchon ex Miquel .......................................................................................... 19
1.2.2 Actinidia melanandra Franchet, Varieties and Synonyms ................... 19
1.2.9 Synonomy of Actinidia fortunatii Finet & Gagnepain,
A. glaucophylla F. Chun, A. henryi Dunn and A. gracilis C. F. Liang ..........21
1.2.10 Actinidia fulvicoma Hance, A. cinerascens C.F. Liang and
A. fulvicoma f. hirsuta (Finet & Gagnepain) C. F. Liang...................... 22
1.2.11 Actinidia hemsleyana var. kengiana (F. P. Metcalf) C. F.Liang and
A. kengiana F. P. Metcalf ................................................................... 22
1.2.12 Actinidia carnosifolia C. Y. Wu, A. carnosifolia var. glaucescens C. F. Liang and A. henryi var. polyodonta Handel-Mazzetti ...................... 22
1.2.13 Actinidia flavofloris H. Z. Jiang ......................................................... 22
1.2.14 Actinidia leptophylla C. Y. Wu, A. maloides H. L. Li and A. maloides
f. cordata C. F. Liang ......................................................................... 22
1.2.15 Actinidia jiangkouensis S. D. Shi & Z. S. Zhang and A. laevissima var. floscula S. D. Shi ......................................................................... 22
1.2.16 Actinidia guilinensis C. F. Liang.........................................................23
1.2.17 Actinidia linguiensis R. G. Li & X. G. Wang and A. wantianensis
R. G. Li & L. Mo ................................................................................. 23
1.2.18 Actinidia longicarpa R. G. Li & M. Y. Liang and A. rubraf ilmenta
R. G. Li & J. W. Li ............................................................................... 23
1.2.19 Actinidia petelotii Diels and A. rudis Dunn ........................................ 23
1.2.20 Actinidia pentapetala R. G. Li & J. W. Li............................................. 23
1.2.21 Actinidia zhejiangensis C.F. Liang and Actinidia persicina R. H. Huang & S. M. Wang ........................................................................ 23
1.2.22 Actinidia vitifolia C.Y. Wu and A. rubus Léveillé................................. 23
1.2.23 Actinidia rubricaulis Dunn and A. fasciculoides C.F. Liang......................... 23
1.2.24 Actinidia ulmifolia C.F. Liang, A. sorbifolia C.F. Liang, A.
stellato-pilosa C.Y. Chang, A. obovata Chun ex C.F. Liang and
A. grandiflora C.F. Liang ................................................................... 23
1.3 Paleopolyploid Origin, Biogeographic Patterns and Reticulate Evolution ...........................................................................23
1.3.1 Paleopolyploid Origin and Evolution of Actinidia .............................. 23
1.3.2 Biogeographic Patterns and Reticulate Evolution .............................. 26
1.4.1 Pollen Size and Morphology .............................................................. 29
1.4.2 Phylogenetic Relationships in Actinidia as Revealed by Pollen Morphology ....................................................................................... 30
1.4.3 Applications of Palynology ................................................................ 31
1.5 Flower Morphology and Sex Variation.............................................34
1.6 Ploidy Variation and Breeding Applications ...................................36
1.6.1 Basic Chromosome Number .............................................................. 36
1.6.2 Ploidy Races and Interploidy Hybridization........................................ 36
1.6.3 Nuclear DNA Content and Flow Cytometry......................................... 41
1.6.4 Ploidy Variation for Kiwifruit Breeding............................................... 42
3 Natural Distribution of Genus Actinidia.......................... 167
3.1 Regional Distribution, Natural Habitats, and Wild Production ......................................................................................... 167
3.1.1 China ............................................................................................... 167
3.1.2 East Asia and Southeast Asia ........................................................... 181
3.2 Vertical Distribution ........................................................................ 183
3.3 Geographic Distribution of Commercially Important Species ........ 183
3.4 Geographically Overlapping Distributions, Natural hybridization and Gene Introgression of Species ................................................... 187
4 Domestication and Commercialization of Actinidia....... 189
4.1 Use of Actinidia Natural Resources in Ancient China .................. 189
4.2 Early Western Introductions of Actinidia Species .......................... 191
4.3 Domestication and Commercialization of A. chinensis var. chinensis and A. chinensis var. deliciosa ........................................ 192
4.3.1 First Botanical Specimens ................................................................192
4.3.2 Cultivation of A. chinensis var. deliciosa Outside China ..................... 193
4.3.3 Commercialization of A. chinensis var. deliciosa ............................... 195
4.3.4 Early Attempts to Cultivate A. chinensis var. chinensis Outside China ............................................................................................... 197
4.4 Domestication and Commercialization of A. chinensis var. chinensis in China ............................................................................ 198
4.4.2 Evaluation of Wild Actinidia Germplasm in China............................. 199
4.4.3 Start of Commercial Cultivation of A. chinensis var. chinensis in China ............................................................................................... 200
4.4.4 Commercialization of A. chinensis var. chinensis Outside China ............................................................................................... 200
4.5 Domestication and Commercialization of A. kolomikta,
A. arguta, A. polygama and A. eriantha......................................... 202
4.6 The World Kiwifruit Industry.......................................................... 203
4.6.1 World Plantings of Kiwifruit............................................................. 203
4.6.2 World Production of Kiwifruit .......................................................... 204
4.6.3 Important Kiwifruit Cultivars in International Trade......................... 204
4.6.4 Segmentation of the Kiwifruit Market ............................................. 206
5 Genetic Improvements and Cultivar Development
of Actinidia ...................................................................................... 209
5.1 Variation in Important Fruit Characteristics of Actinidia........... 209
5.1.1 Fruit Size .......................................................................................... 209
5.1.2 Fruit Shape ...................................................................................... 211
5.1.3 Fruit Hairs and Skin .......................................................................... 211
5.1.4 Fruit Color ........................................................................................215
5.1.10 Other Fruit Characteristics ............................................................. 222
5.2 Progress in the Selection and Breeding of Actinidia .................. 222
5.2.1 Direct Introduction and Selection from the Wild.............................. 222
5.2.2 Selection from Seed Collected in the Wild........................................ 223
5.2.3 Selection from Open Pollination of Cultivars or Named Selections ........................................................................................ 223
6.2.1 Hort16A (Earligold, fruit marketed as Zespri. Gold Kiwifruit) .......... 239
6.2.2 Jintao (Wuzhi 81-1, C6, WIB-C-6, fruit marketed outside China under the names Jingold. and Kiwita) .......................................... 240